Having seen Demetrius act cruelly toward Helena, Oberon orders Puck to spread some of the elixir on the eyelids of the young Athenian man. Instead, Puck accidentally puts the juice on the eyes of Lysander, who then falls in love with Helena. Oberon sees Demetrius still following Hermia and is enraged. When Demetrius decides to go to sleep, Oberon sends Puck to get Helena while he charms Demetrius' eyes. Due to Puck's mistake of putting the juice on Lysander's eyes, both lovers now fight over Helena instead of Hermia. Helena, however, is convinced that her two suitors are mocking her, as neither loved her originally. The four quarrel with each other until Lysander and Demetrius become so enraged that they seek a place to duel each other to the death to prove whose love for Helena is the greatest. Oberon orders Puck to keep Lysander and Demetrius from catching up with one another and to remove the charm from Lysander, so that he goes back to being in love with Hermia.
Meanwhile, a band of six lower-class labourers ("rude mechanicals", as they are described by Puck) have arranged to perform a play about Pyramus and Thisbe for Theseus' wedding and venture into the forest, near Titania's bower, for their rehearsal. Nick Bottom, a stage-struck weaver, is spotted by Puck, who transforms his head into that of a donkey. When Bottom returns for his next lines, the other workmen take one look at him and run screaming in terror. Determined to wait for his friends, he begins to sing to himself. Titania is awakened by Bottom's singing and immediately falls in love with him. She treats him like a nobleman and lavishes him with attention. While in this state of devotion, she encounters Oberon and casually gives him the Indian boy. Having achieved his goals, Oberon releases Titania and orders Puck to remove the ass's head from Bottom. The magical enchantment is removed from Lysander but is allowed to remain on Demetrius, so that he may reciprocate Helena's love.
The fairies then disappear, and Theseus and Hippolyta arrive on the scene, during an early morning hunt. They wake the lovers and, since Demetrius doesn't love Hermia anymore, Theseus over-rules Egeus's demands and arranges a group wedding. The lovers decide that the night's events must have been a dream. After they all exit, Bottom awakes, and he too decides that he must have experienced a dream "past the wit of man." In Athens, Theseus, Hippolyta and the lovers watch the six workmen perform Pyramus and Thisbe. It is ridiculous and badly performed but gives everyone pleasure regardless, and afterward everyone retires to bed. Afterwards, Oberon, Titania, Puck, and other fairies enter, and bless the house and its occupants with good fortune.

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